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Rachid Amara

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Professeur des universités - Universités du Littoral Côte d’Opale

Adresse Contact
MREN Tél : 03 21 99 64 34
32 avenue Foch Fax : 03 21 99 64 01
62930 Wimereux, france Email : Rachid Amara

Thèmes de recherche :

Formation :

Publications internationales à comité de lecture :

- AMARA, R., LAGARDÈRE, F. & DESAUNAY, Y. 1993. Seasonal distribution and duration of the planktonic stage of Dover sole, Solea solea, larvae in the Bay of Biscay : an hypothesis. Journal of Fish Biology. 43 : 17-30.

- AMARA, R., DESAUNAY, Y. & LAGARDERE, F. 1994. Seasonal variation in growth of larval sole Solea solea (L.) and consequences on the success of larval immigration. Netherlands Journal of Sea Research. 32 : 287-298.

- SCIANDRA, A. & AMARA, R. 1994. Effects of nitrogen limitation on growth and nitrite excretion rates of the dinoflagellate Prorocentrum minimum. Marine Ecology Progress Series. 105 : 301-309.

- AMARA, R. & LAGARDERE, F. 1995. Taille et âge au début de la métamorphose chez la sole Solea solea L. du golfe de Gascogne. ICES. Journal of Marine Science. 52 : 247-256.

- AMARA, R. & BODIN, P. 1995. L’environnement trophique méiobenthique en dehors des nourriceries côtières est-il favorable à l’installation des jeunes soles ? Océanologica Acta. 18 (5) :1-12.

- AMARA R ; J.C. POULARD ; F. LAGARDÈRE & Y. DÉSAUNAY.1998. Comparison of the life cycle of two soleidae : Solea solea (L.) and Microchirus variegatus (Donovan) in the Bay of Biscay. Environmental Biology of Fish. 53 : 193-209.

- LAGARDÈRE, F ; R. AMARA & L. JOASSARD.1999. Vertical distribution and feeding activity of metamorphosing sole, Solea solea (L.), before immigration to a coastal nursery the Bay of Vilaine (Northern Bay of Biscay). Environmental Biology of Fish. 56 : 213-228.

- AMARA, R., LAGARDERE, F. & DESAUNAY, Y.& J. MARCHAND. 2000. Metamorphosis and estuarine colonisation in the common sole, Solea solea (L.) : implications for recruitment regulation . Océanologica Acta. 23 : 469-484.

- AMARA R., P. LAFFARGUE., J.M. DEWARUMEZ., C. MARYNIACK, F. LAGARDERE AND C. LUCZAC. 2001. Feeding ecology and growth of O-group flatfishes (sole, dab and plaice) at a nursery ground on the southern North Sea. Journal of Fish Biology. 58 : 788-803.

- AMARA R & C. PAUL. 2003. Seasonal patterns in the fish and epibenthic crustaceans community of an intertidal zone with particular reference to the population dynamics of plaice and brown shrimp. Estuarine Coastal and Shelf Sciences.56 : 809-820.

- AMARA, R. 2003. Seasonal Ichthyodiversity and Growth Patterns of Juvenile Flatfish on a Nursery Ground in the Southern Bight of the North Sea (France). Environmental Biology of Fishes 67(2) : 191-201.

- DE PONTUAL, H ; F. LAGARDERE ; R. AMARA ; M. BOHN & A. OGOR. 2003. Influence of ontogenetic and environmental changes in the otolith microchemistry of juvenile sole (Solea solea). Journal of Sea Research. 50(1/2) : 199-210.

- AMARA R ; K. MAHE ; O. LEPAPE & N. DESROY. 2004. Feeding, growth and distribution of solenette Buglossidium luteum (Risso, 1810) with particular references to its habitat preferences. Journal of Sea Research. 51 : 211-217.

- AMARA, R. 2004. O-group flatfish growth conditions on a nursery ground (bay of Canche, Eastern English Channel). Hydrobiologia, 518, 23-32.

- AMARA, R & R. GALOIS. 2004. Nutritional condition of metamorphosis sole, Solea solea (L.) : spatial and temporal analyses. Journal of Fish Biology, 64 : 1-17.

- HENRY, F ; R. AMARA ; L. COURCOT ; D. LACOUTURE & M.L. BERTHO. 2004. Spatial comparison of heavy metals in fish from the French coast of the Eastern English Channel and Southern bight of the North Sea. Environment International. 30 : 675-683.

- GILLIERS, C ; R. AMARA & J.P. BERGERON. 2004. Quality of flatfish nursery habitats in the Eastern Channel, and southern Bight of the North Sea. Environmental Biology of Fishes. 71 : 189-198.

- KACHER, M. & R. AMARA. 2005. Distribution and growth of O-group juvenile European hake in the Bay of Biscay and Celtic Sea : a spatial and inter-annual analyses. Fisheries Research . 71 : 373-378

- GILLIERS, C., O. LE PAPE, Y. DÉSAUNAY, J. MORIN, R. AMARA (2006) Are growth and density quantitative indicators of essential fish habitat quality ? An application to the common sole (Solea solea L.) nursery grounds. Estuarine, Coastal and Shelf Science, 69, 1-2, 96-106.

- GILLIERS, C ; O. LEPAPE ; Y. DESAUNAY, J.P. BERGERON, N. SCHREIBER & R. AMARA. 2006. Growth and condition of juvenile sole (Solea solea L.) as indicator of habitat quality in coastal and estuarine nurseries in the Bay of Biscay with a focus on sites exposed to the Erika oil spill.. Scientia Marina. 70, 183-192

- AMARA R, MEZIANE T, GILLIERS C, HERMEL G, LAFFARGUE P. 2007. Growth and condition indices in juvenile sole (Solea solea L.) measured to assess the quality of essential fish habitat. Marine Ecology Progress Series 351:209-220

- BACHA, M & R. AMARA. 2007. Les poissons des eaux continentales d’Algérie. Étude de l’ichtyofaune de la Soummam. Cybium.31 : 351-358

- LAFFARGUE P, F. LAGARDERE, A. D. RIJNSDORP, A. FILLON, R. AMARA. 2007. Growth performances of juvenile sole Solea solea under environmental constraints of embayed nursery areas. Aquatic Living Resources, 20 : 213-221

- LE PAPE O., GILLIERS, C., RIOU, P., MORIN J & AMARA, R. 2007. Convergent signs of degradation of both the capacity and the quality of an essential fish habitat : synthesis on the Seine estuary (France), an highly anthropised estuary of which flatfish nursery function is altered. Hydrobiologia. 588 : 225-229

- MAHE, K, R. AMARA, T. BRYCKAERT, M. KACHER, AND J. M. BRYLINSKI. 2007. Ontogenetic and spatial variation in the diet of hake (Merluccius merluccius) in the Bay of Biscay and the Celtic Sea ICES Journal of Marine Sciences 64 : 1210-1219

- SELLESLAGH, J & R. AMARA. (2007) Influence of environmental conditions on macrofauna assemblages in intertidal zone. Cybium 31 (2) : 155-162

- SELLESLAGH, J & R. AMARA. 2008. Inter-season and interannual variations in fish and macrocrustacean community structure on a eastern English Channel sandy beach : influence of environmental factors. Estuarine, Coastal and Shelf Science, 77 : 721-730.

- VINAGRE, C, V. FONSECA, A. MAIA, R. AMARA AND H. CABRAL.2008. Habitat specific growth rates and condition indices for the sympatric solesSolea solea (Linnaeus, 1758) and Solea senegalensis Kaup 1858, in the Tagus estuary, Portugal, based on otolith daily increments and RNA-DNA ratio. J. Appl. Ichthyol. 24 (2008), 163–169

- VINAGRE, C, R. AMARA, A. MAIA, H.N. CABRAL. 2008. Latitudinal comparison of spawning season and growth of 0-group sole, Solea solea (L.). Estuarine, Coastal and Shelf Science . 78 : 521-528.

- SELLESLAGH, J & R. AMARA. 2008. Environmental factors structuring fish composition and assemblages in a small macrotidal estuary (eastern English Channel). Estuarine, Coastal and Shelf Science, 79, 507-517.

- SELLESLAGH J, R. AMARA , P. LAFFARGUE, S. LESOURD , M. LEPAGE, M. GIRARDIN. 2009.. Composition and functioning of three fish estuarine assemblages of the eastern English Channel : a comparison with French estuaries. Estuarine, Coastal and Shelf Science, 81 : 149-159.

- COURRAT A, J. LOBRY, D. NICOLAS, P. LAFFARGUE, R. AMARA, M. LEPAGE, M. GIRARDIN, O. LE PAPE. 2009. Anthropogenic disturbance on nursery function of estuarine areas for marine species. Estuarine, Coastal and Shelf Science. 81 : 179-190.

- AMARA, R , J. SELLESLAGH, G. BILLON, C. MINIER. 2009. Growth and condition of 0-group European flounder, Platichthys flesus as indicator of estuarine habitat quality. Hydrobiologia 67:87-98.

- BACHA M , AMARA, R. 2009. Spatial, temporal and ontogenetic variation in diet of anchovy (Engraulis encrasicolus L) on the Algerian coast (SW Mediterranean). Estuarine coastal and Shelf Sciences 85, 257–264.

- BACHA M, AMARA, R., MOALI, A, BENMANSOUR, N, BRYLINSKI J.M & MAHÉ, K , 2010 - Relationships between age, growth, diet and environmental parameters for anchovy (Engraulis encrasicolus L.) in the Bay of Benisaf (SW Mediterranean, west Algerian coast). Cybium 34, 47-57.

- AMARA R. 2010. Impact de l’anthropisation sur la biodiversité et le fonctionnement des écosystèmes marins. Exemple de la Manche-mer du nord. », VertigO - la revue électronique en sciences de l’environnement, Hors-série 8 | octobre 2010, URL :

- AMARA R. 2011. Impact de la pollution sur les écosystèmes côtiers : exemple de la Manche orientale. VertigO - la revue électronique en sciences de l’environnement, Hors-série 9 | juillet 2011 URL :

- KERAMBRUN, E. SANCHEZ, W., HENRY, F. AND AMARA, R. 2011. Are biochemical biomarker responses related to physiological performance of juvenile sea bass (Dicentrarchus labrax) and turbot (Scophthalmus maximus) caged in a polluted harbour ? Comparative Biochemistry and Physiology Part C : Toxicology & Pharmacology, 154 : 187-195

- BENGUEDDA W , N. DALI YOUCEF AND R. AMARA 2011.Trace metals in sediments, macroalgae and benthic species from the western part of Algerian coast. Journal of Environmental Science and Engineering 15 (12) : 1604-1612.

- SELLESLAGH J ; J. LOBRY , R. AMARA, P. BOET. 2011. Trophic functioning of coastal ecosystems along an anthropogenic pressure gradient : A French case study with emphasis on a small and low impacted estuary. Estuarine, Coastal and Shelf Science, 112 : 73-85.

- KERAMBRUN, E., F. HENRY, P. PERRICHON, L. COURCOT, T. MEZIANE, N. SPILMONT, R. AMARA. 2012. Growth and condition indices of juvenile turbot, Scophthalmus maximus, exposed to contaminated sediments : Effects of metallic and organic compounds. Aquatic Toxicology, 108:130-140.

- SELLESLAGH J., S. LESOURD AND R. AMARA. 2012. Comparison of macrobenthic assemblages of three fish estuarine nurseries and their importance as foraging grounds, Journal of the Marine Biological Association of the United Kingdom, 92 : 85-97.

- KERAMBRUN, E., S. LE FLOCH, H. THOMAS-GUYON, T. MEZIANE, F. HENRY AND R. AMARA. 2012. Responses of juvenile sea bass, Dicentrarchus labrax, exposed to acute concentrations of crude oil, as assessed by molecular and physiological biomarkers. Chemosphere 87, 692-702.

- KERAMBRUN, E. HENRY, F., COURCOT, L., GEVAERT, F., AND AMARA, R. 2012. Biological responses of caged juvenile sea bass (Dicentrarchus labrax) and turbot (Scophtalmus maximus) in a harbour polluted area. Ecological Indicators 19 : 161-171.

- BACHA M , AMARA, R. 2012. Inter-cohort differences in growth, condition and feeding of juvenile anchovy (Engraulis encrasicolus) in the Gulf of Béjaia (Algerian coast, SW Mediterranean) : Implications for recruitment success. Fisheries Research, 129 : 73– 81

- KERAMBRUN, E, F. HENRY, A. MARÉCHAL A., W. SANCHEZ, C. MINIER, I. FILIPUCI AND R. AMARA. 2012. A multibiomarker approach in juvenile turbot, Scophthalmus maximus, exposed to contaminated sediments. Ecotoxicology and Environmental Safety 80, 45-53.

- HENRY F, I. FILIPUCI, G. BILLON, L. COURCOT, E. KERAMBRUN AND R. AMARA. 2012. Metal concentrations, growth and condition indices in European juvenile flounder (Platichthys flesus) relative to sediment contamination levels in four Eastern English Channel estuaries. J. Environ. Monit., 2012, 14, 3211-3220.

- SELLESLAGH J AND AMARA R. 2012. Effect of starvation on condition and growth of juvenile plaice Pleuronectes platessa : nursery habitat quality assessment during the settlement period. Journal of the Marine Biological Association of the United Kingdom, 1-10.

- Kerambrun, E., Henry, F., Cornille, V., Courcot, and Amara R., 2013. A combined measurement of bioaccumulation and condition indices in juvenile European flounders, Platichthys flesus, from European estuaries. Chemosphere. 91 : 498–505

- VINAGRE, C, R. AMARA, H. CABRAL. 2012. Spawning period of Senegal sole, Solea senegalensis, based on juvenile otolith microstructure. Journal of Sea Research 76 (2013) 89–93

- RAMDANE, Z., J.P. TRILLES., K. MAHE ET R. AMARA. 2012. Metazoan ectoparasites of two Teleostei species, Boops boops (L.) and Mullus barbatus barbatus L. from Algerian coast : diversity, parasitological index and impact of parasitism. Cybium (sous presse).

- KERAMBRUN, E., HENRY, F., SANCHEZ, W., AND AMARA, R., 2012. Relationships between biochemical and physiological biomarkers responses measured on juvenile marine fish to environmental chemical contamination. Comparative Biochemistry and Physiology Part A : Molecular & Integrative Physiology, Supplement September 63, 22–23.

- KERAMBRUN, E., AMARA, R ., HENRY, 2013. Effects of food limitation on 9 metal concentration in liver and polycyclic aromaitic hydrocarbon metabolites in bile of juvenile turbot (Scophthalmus maximus) previously exposed to contaminated sediments. Environmental Toxicology and Chemistry, Vol. 32, No. 11, pp. xx–xx, 2013.

- Amara R., I. Filipuci, D. Vincent , F. Goulard, E. Breton. 2013. Do transparent exopolymeric particles (TEP) derived from Phaeocystis globosa bloom impact the physiological performances of European sea bass juveniles. Aquaculture 414–415 (2013) 149–154.

Ouvrages individuels et collectifs :

- LAGARDERE, F., CHAUMILLON, G., AMARA, R., HEINEMAN, G. & LAGO, J.M. 1995. Examination of otolith morphology and microstructure using Laser Scanning Microscopy, p. 7-27 . In : Secor, D.H., J.M. Dean and S.E. Campana. Recent developments in fish otolith research. Belle W. Baruch Library in Marine Science N°. 19. Univ. South Carolina Press, Columbia, South Carolina. 733 pp.

- MINIER, C & R. AMARA. 2009. From pollution to altered fish biological and physiological performances ; the case of flatfish in the Seine estuary. In ‘Environmental assessment of estuarine ecosystems : a case study’. Eds : C. Amiard-Triquet and P.S. Rainbow, CRCPress/Taylor and Francis. 344 pp.